Galium divaricatum

Pourr. ex Lam. (1788)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Rubiaceae Genus: Galium

bedstraw

Description

Key Characters:

Growth Form: Diffuse, slender, branched annual herbs.

Stems: Stems 0.5–3 dm long, slightly scabrid, at least in lower portions.

Roots:

Leaves: Leaves simple. Whorled, 6–8 per node. Blades narrowly lanceolate to linear, 4–10 mm long, 0.3–1.5(–2) mm wide. Upper surfaces usually slightly hispid. Margins slightly revolute, margins and midrib antrorsely scabrid. Stipules leaf-like.

Flowers: Flowers in delicate, lax inflorescences, peduncles 5–20 mm long, pedicels filiform, 0.5–3 mm long, deflexed after anthesis. Flowers insect-pollinated, bisexual (perfect) or rarely unisexual, often heterostylous. Sepals obsolete. Corolla yellowish red, 0.5–1 mm in diameter, (3)4-lobed. Stamens 3–4 and alternate with the corolla lobes; filaments usually inserted on the corolla tube or throat; anthers dithecal, opening by longitudinal slits. Ovary inferior, occasionally partly so, or very rarely superior, 2-lobed, 2-celled; ovules 1 per cell; styles 2, terminal, slender; stigmas capitellate, dry or occasionally wet.

Fruit: Fruit didymous; dry or fleshy; 0.5–0.7 mm long; consisting of 2 indehiscent cocci usually separating at maturity; finely papillose. Seeds with well-developed oily endosperm; or endosperm occasionally scanty or absent.

Ploidy:

Habitat: Sparingly naturalized on damp cliffs.

Elevation Range: 970–980 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Specimens

Bibliography

Name Published In: Encycl. 2: 580 (1788)

Other References

Wagner et al. 1990:1131 (EM); Wagner & Herbst 1995:25 (H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Galium divaricatum Pourr. ex Lam. Koloa District: National Tropical Botanical Garden in Lawai Valley Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Murica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr F Kauai HAPI 8/5/2003
2 Galium divaricatum Pourr. ex Lam. Haleakala National Park, Leileiwi Overlook, by parking lot, other side of walk where it meets road. 2296279N, 788413E. Other side of walk where it meets road, in crack. Subalpine shrubland/roadside with Sophora chrysophylla and Coprosma montana. PRESERVED_SPECIMEN Starr, F. Starr-13081-05 Maui BISH 8/1/2013
3 Galium divaricatum Pourr. ex Lam. Ocean View, HI, USA PRESERVED_SPECIMEN F. Starr, K. Starr & P. Bily 030805-1 Maui HAPI 8/5/2003
4 Galium divaricatum Pourr. ex Lam. waihoi spring, olinda Growing on a damp cliff face in partial sun. Very loose, twining in low grass and ferns. PRESERVED_SPECIMEN hobdy, r.w. collector number: 1087 Maui BISH 5/13/1981
5 Galium divaricatum Pourr. ex Lam. east maui, pohakuokala gulch Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: Mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Myrica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN starr, f. starr, k.; p.bily collector number: 030805-1 Maui BISH 8/5/2003
6 Galium divaricatum Pourr. ex Lam. East Maui, Pohakuokala Gulch Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: Mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Myrica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr, F. 030805-1 Maui BISH 8/5/2003
7 Galium divaricatum Pourr. ex Lam. Pohakuokala Gulch. Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Murica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr F Starr, F 30805- 1 L 8/5/2003
8 Galium divaricatum Pourr. ex Lam. E Maui, Pohakuokala Gulch PRESERVED_SPECIMEN Starr, F.; Starr, K.; Bily, P. 030805-1 Maui US 8/5/2003
9 Galium divaricatum Pourr. ex Lam. E. Maui, Makaeha, Kailua Gulch above Waihou Springs. PRESERVED_SPECIMEN Hank Oppenheimer H20131 Maui PTBG 2/24/2001
10 Galium divaricatum Pourr. ex Lam. E. Maui, Pohakuokala Gulch. Mesic forest, basalt (Kula) plastering lava flow. PRESERVED_SPECIMEN Forest Starr Starr-030805-1 Maui PTBG 8/5/2003
11 Galium divaricatum Pourr. ex Lam. E Maui, Pohakuokala Gulch PRESERVED_SPECIMEN F. Starr, K. Starr & P. Bily 030805-1 Maui US 8/5/2003
12 Galium divaricatum Pourr. ex Lam. e maui, pohakuokala gulch PRESERVED_SPECIMEN f. starr, k. starr & p. bily 030805-1 Maui US 8/5/2003
13 Galium divaricatum Pourr. ex Lam. East Maui, Haleakala, Pohakuokala Gulch PRESERVED_SPECIMEN Herbst, D.R. Collector Number: 9706 Maui BISH 10/20/1994
14 Galium divaricatum Pourr. ex Lam. Waihoi Spring, Olinda Growing on a damp cliff face in partial sun. Very loose, twining in low grass and ferns. PRESERVED_SPECIMEN Hobdy, R.W. 1087 Maui BISH 5/13/1981
15 Galium divaricatum Pourr. ex Lam. east maui, haleakala, pohakuokala gulch PRESERVED_SPECIMEN herbst, d.r. collector number: 9706 Maui BISH 10/20/1994
16 Galium divaricatum Pourr. ex Lam. East Maui, Makawao District, Makaeha, Kailua Gulch above Waihou Springs In pasture. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H20131 Maui BISH 2/24/2001
17 Galium divaricatum Pourr. ex Lam. East Maui, Makawao District, Makaeha, Kailua Gulch above Waihou Springs In pasture. PRESERVED_SPECIMEN Oppenheimer, H.L. H20131 Maui BISH 2/24/2001
18 Galium divaricatum Pourr. ex Lam. east maui, makawao district, makaeha, kailua gulch above waihou springs In pasture. PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h20131 Maui BISH 2/24/2001
19 Galium divaricatum Pourr. ex Lam. East Maui, Pohakuokala Gulch Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: Mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Myrica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr, F. Starr, K.; P.Bily Collector Number: 030805-1 Maui BISH 8/5/2003
20 Galium divaricatum Pourr. ex Lam. East Maui, Haleakala, Pohakuokala Gulch PRESERVED_SPECIMEN Herbst, D.R. 9706 Maui BISH 10/20/1994
21 Galium divaricatum Pourr. ex Lam. Koloa District: National Tropical Botanical Garden in Lawai Valley Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Murica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr F Kauai HAPI 8/5/2003
22 Galium divaricatum Pourr. ex Lam. Haleakala National Park, Leileiwi Overlook, by parking lot, other side of walk where it meets road. 2296279N, 788413E. Other side of walk where it meets road, in crack. Subalpine shrubland/roadside with Sophora chrysophylla and Coprosma montana. PRESERVED_SPECIMEN Starr, F. Starr-13081-05 Maui BISH 8/1/2013
23 Galium divaricatum Pourr. ex Lam. Ocean View, HI, USA PRESERVED_SPECIMEN F. Starr, K. Starr & P. Bily 030805-1 Maui HAPI 8/5/2003
24 Galium divaricatum Pourr. ex Lam. waihoi spring, olinda Growing on a damp cliff face in partial sun. Very loose, twining in low grass and ferns. PRESERVED_SPECIMEN hobdy, r.w. collector number: 1087 Maui BISH 5/13/1981
25 Galium divaricatum Pourr. ex Lam. east maui, pohakuokala gulch Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: Mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Myrica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN starr, f. starr, k.; p.bily collector number: 030805-1 Maui BISH 8/5/2003
26 Galium divaricatum Pourr. ex Lam. East Maui, Pohakuokala Gulch Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: Mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Myrica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr, F. 030805-1 Maui BISH 8/5/2003
27 Galium divaricatum Pourr. ex Lam. Pohakuokala Gulch. Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Murica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr F Starr, F 30805- 1 L 8/5/2003
28 Galium divaricatum Pourr. ex Lam. E Maui, Pohakuokala Gulch PRESERVED_SPECIMEN Starr, F.; Starr, K.; Bily, P. 030805-1 Maui US 8/5/2003
29 Galium divaricatum Pourr. ex Lam. E. Maui, Makaeha, Kailua Gulch above Waihou Springs. PRESERVED_SPECIMEN Hank Oppenheimer H20131 Maui PTBG 2/24/2001
30 Galium divaricatum Pourr. ex Lam. E. Maui, Pohakuokala Gulch. Mesic forest, basalt (Kula) plastering lava flow. PRESERVED_SPECIMEN Forest Starr Starr-030805-1 Maui PTBG 8/5/2003
31 Galium divaricatum Pourr. ex Lam. E Maui, Pohakuokala Gulch PRESERVED_SPECIMEN F. Starr, K. Starr & P. Bily 030805-1 Maui US 8/5/2003
32 Galium divaricatum Pourr. ex Lam. e maui, pohakuokala gulch PRESERVED_SPECIMEN f. starr, k. starr & p. bily 030805-1 Maui US 8/5/2003
33 Galium divaricatum Pourr. ex Lam. East Maui, Haleakala, Pohakuokala Gulch PRESERVED_SPECIMEN Herbst, D.R. Collector Number: 9706 Maui BISH 10/20/1994
34 Galium divaricatum Pourr. ex Lam. Waihoi Spring, Olinda Growing on a damp cliff face in partial sun. Very loose, twining in low grass and ferns. PRESERVED_SPECIMEN Hobdy, R.W. 1087 Maui BISH 5/13/1981
35 Galium divaricatum Pourr. ex Lam. east maui, haleakala, pohakuokala gulch PRESERVED_SPECIMEN herbst, d.r. collector number: 9706 Maui BISH 10/20/1994
36 Galium divaricatum Pourr. ex Lam. East Maui, Makawao District, Makaeha, Kailua Gulch above Waihou Springs In pasture. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H20131 Maui BISH 2/24/2001
37 Galium divaricatum Pourr. ex Lam. East Maui, Makawao District, Makaeha, Kailua Gulch above Waihou Springs In pasture. PRESERVED_SPECIMEN Oppenheimer, H.L. H20131 Maui BISH 2/24/2001
38 Galium divaricatum Pourr. ex Lam. east maui, makawao district, makaeha, kailua gulch above waihou springs In pasture. PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h20131 Maui BISH 2/24/2001
39 Galium divaricatum Pourr. ex Lam. East Maui, Pohakuokala Gulch Mesic forest, basalt (Kula) plastering lava flow. Associated vegetation: Mix of natives (Acacia koa, Sadleria) and weeds (Hypericum canariense, Myrica faya, Erigeron karvinskianus). PRESERVED_SPECIMEN Starr, F. Starr, K.; P.Bily Collector Number: 030805-1 Maui BISH 8/5/2003
40 Galium divaricatum Pourr. ex Lam. East Maui, Haleakala, Pohakuokala Gulch PRESERVED_SPECIMEN Herbst, D.R. 9706 Maui BISH 10/20/1994